Manny (voice)

Ray Romano

Acting

Sid (voice)

John Leguizamo

Acting

Diego (voice)

Denis Leary

Acting

Ellie (voice)

Queen Latifah

Acting

Crash (voice)

Seann William Scott

Acting

Eddie (voice)

Josh Peck

Acting

Fast Tony (voice)

Jay Leno

Acting

Lone Gunslinger Vulture (voice)

Will Arnett

Acting

Scrat (voice)

Chris Wedge

Acting

Dung Beetle Dad (voice)

Peter Ackerman

Writing

Glypto Boy Billy / Beaver Girl (voice)

Caitlin Rose Anderson

Acting

Rhino Boy / Beaver Boy (voice)

Connor Anderson

Acting

Mr. Start (voice)

Joseph Bologna

Acting

Elk Boy (voice)

Jack Crocicchia

Acting

Condor Chick (voice)

Peter de Sève

Visual Effects

Additional Voices (voice)

Ariel Winter

Acting

Female Mini Sloth (voice)

Clea Lewis

Acting

Aardvark Dad (voice)

Stephen Root

Acting

Start Girl (voice)

Nicole DeFelice

Acting

Diatryma Mom (voice)

Debi Derryberry

Acting

Start Dad (voice)

Marshall Efron

Acting

Stu / Male Ox (voice)

Tom Fahn

Acting

Molehog Grandpa / Shovelmouth Male (voice)

Jason Fricchione

Acting

Beaver Dad (voice)

George Jacobs

Acting

Glyptodon (voice)

Brian Scott McFadden

Acting

Shovelmouth Boy (voice)

Jansen Panettiere

Acting

Start Boy 1 (voice)

Gregory Romano

Acting

Start Boy 2 (voice)

Matt Romano

Acting

Dodo (voice)

Carlos Saldanha

Directing

Molehog Kid 1 (voice)

Manoela Scarpa Saldanha

Acting

Molehog Kid 2 (voice)

Sofia Scarpa Saldanha

Acting

Diego's Bird Girl (voice)

Reyna Shaskan

Acting

Freaky Male (voice)

James Sie

Acting

Aardvark Mom (voice)

Cindy Slattery

Acting

Female Ox (voice)

Mindy Sterling

Acting

Mrs. Start (voice)

Renée Taylor

Acting

Cholly (voice)

Alan Tudyk

Acting

Additional Voices (voice)

Claudia Besso

Acting

Additional Voices (voice)

Madeleine Martin

Acting

Additional Voices (voice)

Sherry Lynn

Acting

Additional Voices (voice)

Mickie McGowan

Acting

Additional Voices (voice)

Emily Anderson

Acting

Additional Voices (voice)

Jack Angel

Acting

Additional Voices (voice)

Shane Baumel

Acting

Additional Voices (voice)

Bob Bergen

Acting

Additional Voices (voice)

Kyle J. Bogert

Acting

Additional Voices (voice)

Rodger Bumpass

Acting

Additional Voices (voice)

Paul Butcher

Acting

Additional Voices (voice)

Blanca Camacho

Acting

Additional Voices (voice)

John Cygan

Acting

Additional Voices (voice)

Jennifer Darling

Acting

Additional Voices (voice)

Olivia DeLaurentis

Acting

Additional Voices (voice)

Harrison Fahn

Acting

Additional Voices (voice)

Bill Farmer

Acting

Additional Voices (voice)

Crispin Freeman

Acting

Additional Voices (voice)

Tim Kirkpatrick

Acting

Additional Voices (voice)

David H. Kramer

Production

Additional Voices (voice)

Danny Mann

Acting

Additional Voices (voice)

Laura Marano

Acting

Additional Voices (voice)

Matt McCarthy

Acting

Additional Voices (voice)

Laraine Newman

Acting

Additional Voices (voice)

Jan Rabson

Acting

Additional Voices (voice)

Zack Shada

Acting

Additional Voices (voice)

Thomas Sharkey

Acting

Additional Voices (voice)

Ross Simanteris

Acting

Additional Voices (voice)

Rachel Stand

Acting

Additional Voices (voice)

Spencer Lacey Ganus

Acting

Additional Voices (voice)

Jess Harnell

Acting

James (voice)

Alex Sullivan

Acting

Traffic Vulture

James Godwin

Acting